Twenty five new species and five new subspecies of Scholtzia are described: S. bellairsiorum Rye, S. brevistylis Rye, S. brevistylis subsp. prowaka Rye, S. calcicola Rye, S. chapmanii Trudgen ex Rye, S. cordata Trudgen ex Rye, S. corrugata Rye, S. denticulata Rye, S. halophila Rye, S. halophila subsp. meridionalis Rye, S. halophila subsp. mortlockensis Rye, S. inaequalis Rye, S. laciniata Rye, S. longipedata Rye, S. longipedata subsp. procera Rye, S. multiflora Rye, S. oleosa Rye, S. peltigera Rye, S. pentamera Rye, S. pentamera subsp. collina Rye, S. prostrata Rye, S. quindecim Rye, S. recurva Rye, S. subsessilis Rye, S. tenuissima Rye, S. thinicola Rye, S. trilocularis Rye, S. truncata Rye, S. uniflora Rye and S. uniovulata Rye. Scholtzia sp. Shark Bay (M.E. Trudgen 7429) is reduced to synonymy under S. capitata Benth. and most of the other phrase names in the genus are treated as synonyms of the above new taxa. A key is given to aid the identification of approximately 40 species and subspecies. Most taxa have conservation priority.
A revision of the tiurndins (Tribonanthes, Haemodoraceae)
The tiurndins (Tribonanthes Endl.) comprise a small taxonomically challenging genus of geophytes endemic to the Southwest Australian Floristic Region. Six named and one undescribed species of Tribonanthes are recognised in recent literature. Tribonanthes poses the most difficult taxonomic problems remaining for the Haemodoraceae in Western Australia. It is a genus where herbarium studies alone are insufficient to delineate taxa. A knowledge of variation in wild living material has proved essential to resolve taxonomic difficulties in the tiurndins. This revision recognises and comprehensively illustrates living plants of the 12 recognised species, four of which are new (T. elongata E.J.Hickman & Hopper, T. keigheryi E.J.Hickman & Hopper, T. monantha E.J.Hickman & Hopper and T. porphyrea E.J.Hickman & Hopper). Tribonanthes variabilis Lindl. and T. uniflora Lindl. are reinstated. Morphological and molecular data also support the recognition of three new subgenera: subg. Tribonanthes, subg. Salina E.J.Hickman & Hopper and subg. Boya E.J.Hickman & Hopper. Botanical illustration, especially of internal floral morphology, has revealed several previously unrecorded characters that help diagnose the species of Tribonanthes.
A taxonomic review of the Styphelia xerophylla group (Ericaceae: Epacridoideae: Styphelieae)
HISLOP, M. AND PUENTE-LELIÈVRE, C., Nuytsia30: 155–175 (2019)
Species belonging to the Western Australian S. xerophylla (DC.) F.Muell. group (or Group IX) are described and illustrated. The treatment includes two species published in the nineteenth century, S. xerophylla and S. stomarrhena (Sond.) Sleumer, together with four new species, S. angustiflora Hislop & Puente-Lel., S. cernua Hislop & Puente-Lel., S. disjuncta Hislop & Puente-Lel. and S. sulcata Hislop & Puente-Lel. A seventh member of the group, S. longissima Hislop & Puente-Lel., was described in a recent paper and is not treated again here. The morphological attributes of the group are outlined and a key to species provided.
Elionurus tylophorus (Poaceae: Paniceae: Andropogoneae), a new species from the Kimberley region of Western Australia
BARRETT, M.D. AND HANDASYDE, T., Nuytsia30: 177–194 (2019)
A new annual species, Elionurus tylophorus M.D.Barrett & T.Handasyde is described in the genus Elionurus Willd. of the grass tribe Andropogoneae. It differs from all previously described Elionurus species in having keels of the lower fertile glumes ornamented with projections, these often terminated by minute solitary bristles, rather than regular cilia lacking projections, or with projections terminated by hair-tufts in other species. It differs further from E. citreus (R.Br.) Benth. and E. purpureus E.J.Thomps., the only other Australian Elionurus, in having strongly developed fertile glume nerves, rachilla longer and lacking a wing, and presence of lemmas and anthers in the pedicellate spikelet. Elionurus tylophorus is currently known from a single location in the King Leopold Ranges in the central Kimberley region, and has conservation significance. A primary division of Elionurus into two clades is hypothesised based on a preliminary chloroplast gene phylogeny and two morphological synapomorphies, presence of tufted glume hairs and oil streaks.
A key to the species of Thomasia (Malvaceae: Byttnerioideae)
WILKINS, C.F. AND SHEPHERD, K.A., Nuytsia30: 195–202 (2019)
Banksia densa A.R.Mast & K.R.Thiele has until now been regarded as a widespread species in the Western Australian wheatbelt, comprising a widespread nominate variety and a geographically more restricted variety in the southern part of its range (var. parva (A.S.George) A.R.Mast & K.R.Thiele). An informally recognised morphological variant, which matches the type of the species, is geographically restricted to an area on the edge of the species range in the immediate vicinity of Corrigin. Morphological assessment shows that the Corrigin form differs significantly in leaf morphology from the remainder of the species and comprises the true B. densa. The widespread taxon previously referred to as B. densa var. densa, and the southern var. parva, are here raised to species rank as B. zygocephala K.R.Thiele sp. nov. and B. parva (A.S.George) K.R.Thiele comb. et stat. nov. respectively.
One new species of Eremophila R.Br., E. waitii Chinnock, is described and one subspecies of E. glabra (R.Br.) Ostenf., E.glabra subsp. verrucosa Chinnock, is raised to species level and recognised herein as E. viridissima Chinnock.
Triodia veniciae (Poaceae), a new species from the Pilbara region, Western Australia
The Australian hummock grass genus Triodia R.Br. is currently undergoing taxonomic revision following increased botanical survey in remote Australia. This paper describes a recently discovered species ‘soft’ Triodia from the Pilbara region, T. veniciae M.D.Barrett. The lemma lobes of the new species are narrow and produced into a short awnlet, intermediate between the old, and now united, generic concepts of Plectrachne Henrard with awned lemmas and Triodia with lobed lemmas. Triodia veniciae M.D.Barrett is only known from rocky shale slopes of the eastern Chichester Range, and has priority conservation status. Diagnostic character combinations for T. veniciae are given to allow discrimination from all other Triodia.
Styphelia quartzitica and S. rectiloba (Ericaceae: Epacridoideae: Styphelieae), two new, morphologically anomalous species of restricted distribution
Two unique indehiscent species of Calandrinia Kunth. from Western Australia are described and mapped and their affinities are discussed with regards to a recent phylogeny for Australian Calandrinia.
Elevation of Eucalyptus gardneri subsp. ravensthorpensis, and notes on relationships between obligate-seeder and resprouter members of subseries Levispermae (Myrtaceae)
GOSPER, C.,HOPLEY, T.,BYRNE, M.,HOPPER, STEPHEN D.,PROBER, S.M. AND YATES, C.J., Nuytsia30: 247–252 (2019)
This treatment deals with five annual Western Australian species of Hydrocotyle L. and is based on the outcomes of a recent molecular phylogenetic study combined with a re-evaluation of schizocarp morphology. Hydrocotyle perplexa A.J.Perkins is newly named, H. intertexta R.Br ex A.Rich. is reinstated, and the three other species covered are H. glochidiata Benth., H. scutellifera Benth. and H. tetragonocarpa Bunge. Three taxa are placed in synonymy: H. blepharocarpa F.Muell. under H. scutellifera, H. pilifera Turcz. under H. intertexta and H. hispidula var. tenella Benth. under H. perplexa. Lectotypes are selected for five of these names, and full descriptions, distribution maps and images are provided for all five species. Details of the history of recognition of these species, and a series of misapplications of names, are also given.
Update on generic and specific nomenclature in Paracaleana (Drakaeinae), Caladeniinae and a new name in Caladenia (Orchidaceae)
HOPPER, STEPHEN D. AND BROWN, A.P., Nuytsia30: 279–285 (2019)
Accepting guidance on stability of names from the Preamble to successive International Codes of Botanical Nomenclature, and in the interests of minimizing name changes consistent with present scientific evidence, we propose that the Australian orchid genus Paracaleana should be maintained as a distinct genus from Caleana, and that Caladenia remains best recognised as distinct from Cyanicula, Ericksonella, Pheladenia, Glossodia and Elythranthera. Recent proposals to lump these genera unnecessarily creates new names in the absence of compelling new scientific evidence to do so. A new name, Caladenia varians Hopper & A.P.Br., is erected to replace C. vulgata Hopper & A.P.Br. The type of C. vulgata matches Caladenia incensum Hopper & A.P.Br., and the former species is therefore rendered synonymous with the latter.
Paracaleana ferricola (Orchidaceae), a new, rare species from the south-west of Western Australia
BROWN, A.P. AND BROCKMAN, G., Nuytsia30: 287–289 (2019)
Hibbertia spicata F.Muell. and H. polystachya Benth. are widespread in south-western Western Australia, where they are unique in Hibbertia Andrews in having flowers in a cincinnus rather than solitary. The boundary between these two species has been problematic, as they mostly differ in the composition and density of indumentum on their sepals and leaves. Close study has revealed that H. polystachya is widespread and taxonomically relatively straightforward, while H. spicata sens. lat. comprises six distinctive, readily resolvable species (including H. leptotheca (J.R.Wheeler) K.R.Thiele comb. et stat. nov., previously H. spicata subsp. leptotheca J.R.Wheeler) differing in leaf and sepal indumentum, number of flowers in the cincinnus, floral bract shapes, stamen number, and number and arrangement of staminodes. New species described here are H. capensis K.R.Thiele sp. nov., H. asterella K.R.Thiele sp. nov. and H. subglabra K.R.Thiele sp. nov. The new name H. prolata K.R.Thiele is provided for a widespread taxon based on Hemistephus linearis J.Drumm. ex Harv. With these species removed, H. spicata is recircumscribed and is now more restricted in distribution.
A new name, clarification of synonymy, and a new subspecies for Isopogon (Proteaceae) in Western Australia
RYE, B.L. AND MACFARLANE, T.D., Nuytsia30: 309–316 (2019)
Isopogon drummondii Benth. nom. illeg. is lectotypified and replaced by the new name I. autumnalis Rye & T.Macfarlane while I. drummondii Hügel ex Jacques is recognised as a probable synonym of I. sphaerocephalus Lindl. The new subspecies I. sphaerocephalus subsp. lesueurensis Rye is described.