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The journal of the Western Australian Herbarium


Volumes 1–14 are available for download via the Biodiversity Heritage Library’s (BHL) page for Nuytsia.

Displaying records 1–20 of 49

New species and notes on central Australian Goodenia (Goodeniaceae)

ALBRECHT, D.E., Nuytsia 15 (1): 1–9 (2002)

Goodenia halophila Albr. sp. nov. and G. cylindrocarpa Albr. sp. nov. are described, with notes on their distribution, conservation status, habitat and relationships with other species of Goodenia Sm. Notes are also provided on the Northern Territory populations of G. angustifolia Carolin and G. glandulosa K.Krause, and on all populations of G. anfracta J.M.Black and G. maideniana W.Fitzg. Further studies are needed to establish whether G. anfracta, which currently has conservation priority in Western Australia, should be reduced to synonymy under G. maideniana.

A revision of Byblis (Byblidaceae) in south-western Australia

CONRAN, J.G., Nuytsia 15 (1): 11–19 (2002)

The south-western Australian populations of Byblis (Byblidaceae) are revised. A new species, B. lamellata Conran & Lowrie, is recognised on habit, ecology and seed morphology and named for the populations between Arrowsmith River and Cataby. Byblis gigantea Lindl. and its synonym B. lindleyana Planch. are lectotypified. A generic description and a table of distinguishing features for all species is given. A key and distribution map are provided for the two species from the south-west of Western Australia.

Conostephium magnum (Epacridaceae), a new species from Western Australia

CRANFIELD, R.J., Nuytsia 15 (1): 21–25 (2002)

A new species of Epacridaceae endemic to the South West Botanical Province of Western Australia, Conostephium magnum Cranfield, is described, illustrated and mapped. It is related to C. minus Lindl. and C. pendulum Benth.

A comparison of the leaf anatomy of Ficus subpuberula, F. atricha, and F. brachypoda (Moraceae: Urostigma sect. Malvanthera)

DIXON, D.J., Nuytsia 15 (1): 27–32 (2002)

The leaf anatomy of Ficus subpuberula Corner, F. atricha D.J.Dixon, and F. brachypoda (Miq.) Miq. are compared in order to facilitate identification of these partly sympatric species. Ficus subpuberula was found to possess distinctly thinner (267.6 ± 4.3 µm), isobilateral leaves compared to the much thicker (313.3 ± 12.1 µm and 425.6 ± 17.8 µm), dorsiventral leaves of F. atricha and F. brachypoda respectively. Tanniniferous cells were present in F. atricha and F. brachypoda, but absent in F. subpuberula. The upper epidermis and the palisade parenchyma of F. brachypoda were about twice as thick as in F. atricha.

A new species of Blennospora (Asteraceae: Gnaphalieae) from the Swan and Scott Coastal Plains of Western Australia

KEIGHERY, G.J., Nuytsia 15 (1): 33–36 (2002)

A new species of Blennospora A. Gray, B. doliiformis Keighery, is described from the Swan and Scott Coastal Plains. The species occurs on clay soils in fresh winter-wet sites. A key is given to the three species now recognised in the genus.

A review of the genus Pileanthus (Myrtaceae)

KEIGHERY, G.J., Nuytsia 15 (1): 37–51 (2002)

Pileanthus Labill., a small genus of floriferous shrubs endemic to southern Western Australia, is reviewed. Eight species are recognised, one of which has two subspecies. Four new species (Pileanthus auranticus Keighery, P. bellus Keighery, P. rubronitidus Keighery and P. septentrionalis Keighery) and one new subspecies (P. peduncularis subsp. piliferus Keighery) are described, and P. vernicosus F. Muell. is reinstated.

Two new species of Comesperma (Polygalaceae) from Western Australia

KEIGHERY, G.J., Nuytsia 15 (1): 53–57 (2002)

Two new species of Comesperma Labill. from the south-west of Western Australia, C. calcicola Keighery and C. griffinii Keighery, are described and illustrated. Both species are regarded as poorly collected and in need of further survey to determine their conservation status. The great diversity of life forms found in south-western members of the genus is outlined.

Drosera pedicellaris (Droseraceae), a new species from south-west Western Australia

LOWRIE, A., Nuytsia 15 (1): 59–62 (2002)

A new species, Drosera pedicellaris Lowrie, is described and illustrated. It is related to D. parvula Planchon and is known from only two localities in the south-west of Western Australia.

Eucalyptus calyerup (Myrtaceae), a new species of possible hybrid origin from south-western Australia

MCQUOID, N.K. AND HOPPER, STEPHEN D., Nuytsia 15 (1): 63–68 (2002)

Eucalyptus calyerup McQuoid & Hopper is described and illustrated. It appears to be of possible hybrid origin, but morphologically stable, with the putative parents being Eucalyptus occidentalis Endl. and E. platypus Hook. subsp. platypus. It is endemic to the Calyerup Rocks area near Jerramungup in Western Australia and has aroused much interest from landcare groups. It is rare in the wild and in need of monitoring.

A revision of Eucalyptus series Calcicolae (Myrtaceae) from the south coast of Western Australia

NICOLLE, D., Nuytsia 15 (1): 69–76 (2002)

Eucalyptus series Calcicolae Brooker consists of four closely related taxa, two of which are newly described. E. calcicola Brooker subsp. calcicola is restricted to the Hamelin Bay area on the Leeuwin Coast, and subsp_. unita_ Nicolle subsp. nov. occurs in scattered populations from west of Denmark eastwards towards Bremer Bay. E. ligulata Brooker subsp. ligulata occurs from Cape Le Grand to Cape Arid, and subsp. stirlingica Nicolle subsp. nov. is restricted to the Stirling Range. A key and distribution map for the taxa of Eucalyptus ser. Calcicolae are presented.

Two new species of silver mallet (Eucalyptus - Myrtaceae) of very restricted distribution in south-western Western Australia

NICOLLE, D., Nuytsia 15 (1): 77–83 (2002)

Eucalyptus rugulata Nicolle sp. nov. is described from the South Ironcap-Hatter Hill area north-east of Lake King, distinguished within the silver mallets by its broad and thick adult leaves and the large and robust buds and fruits. E. purpurata Nicolle sp. nov. is described from a single population near Bandalup Hill east of Ravensthorpe. It is distinguished from E. argyphea L.A.S. Johnson & K.D. Hill by the conspicuous red-purple new growth and the smaller buds and fruits. Both new species are of restricted distribution and E. purpurata, in particular, is considered to be at risk. A key and distribution map for the silver mallets are presented.

One new Banksia and two new Grevillea species (Proteaceae: Grevilleoideae) from Western Australia

OLDE, PETER M. AND MARRIOTT, NEIL R., Nuytsia 15 (1): 85–99 (2002)

Banksia rosserae P. Olde & N. Marriott, a new species of Banksia R. Br., Grevillea kirkalocka P. Olde & N. Marriott and G. squiresae P. Olde & N. Marriott, two new species of Grevillea R. Br. (Proteaceae: Grevilleoideae) are described. Their affinities are discussed and keys are provided. All of the new species have conservation priority.

A revision of the south-western Australian species of Micromyrtus (Myrtaceae) with five antisepalous ribs on the hypanthium

RYE, B.L., Nuytsia 15 (1): 101–122 (2002)

Among the species currently included in Micromyrtus Benth., two main categories are distinguished here on the basis of hypanthium ribbing, one characterised by having antipetalous ribs and usually also antisepalous ones, the other by having antisepalous ribs but no antipetalous ones. Most species belong to the latter category, including the lectotype selected here for the genus, Micromyrtus drummondii Benth. nom. superfl. [= M. obovata (Turcz.) J.W. Green]. A key is given to the 14 Western Australian members of this group, and the eight south-western species, including five new species, are revised. Micromyrtus erichsenii Hemsl. is reinstated and the new species M. monotaxis Rye, M. ninghanensis Rye, M. papillosa J.W. Green ex Rye, M. rogeri J.W. Green ex Rye and M. uniovula Rye are illustrated. Three of these new species have an ovule number of one, lower than any previously recorded for the genus, and four of them have conservation priority.

Scaevola ballajupensis (Goodeniaceae), a new species from the Jarrah forest of south-western Australia

SAGE, L.W., Nuytsia 15 (1): 123–126 (2002)

The new species Scaevola ballajupensis L.W. Sage, from the south-western Australian Jarrah forest, is described and photographically illustrated. The species is known only from one locality on private property and therefore has a high conservation priority. An amendment to the key to Scaevola species in "Flora of Australia" is made to include the new taxon and a table showing how it differs from its closest relatives is provided.

A revision of Hibbertia depressa and its allies (Dilleniaceae) from Western Australia

WHEELER, J.R., Nuytsia 15 (1): 127–138 (2002)

A small group of closely related taxa, which form part of section Candollea Gilg. in the genus Hibbertia Andr., are revised. Descriptions, illustrations and distribution maps are presented along with a key, including two new taxa, Hibbertia fitzgeraldensis J.R. Wheeler and Hibbertia priceana J.R. Wheeler. Both new species are restricted in distribution and Hibbertia priceana appears to be at risk. The lectotypification of Hibbertia desmophylla (Benth.) F. Muell. is also made.

Miscellaneous new species of Hibbertia (Dilleniaceae) from the wheatbelt and pastoral regions of Western Australia

WHEELER, J.R., Nuytsia 15 (1): 139–152 (2002)

The new taxa, Hibbertia ancistrophylla J.R. Wheeler, H. ancistrotricha J.R. Wheeler, H. avonensis J.R. Wheeler, H. lepidocalyx J.R. Wheeler, H. lepidocalyx subsp. tuberculata J.R. Wheeler and H. stenophylla J.R. Wheeler are described, mapped and illustrated. All belong to either section Pleurandra (Labill.) Benth. or section Hemipleurandra Benth. They are recorded from the Avon Wheatbelt, Jarrah Forest and Mallee regions of the South West Botanical Province and the Coolgardie, Murchison and Yalgoo regions of the Eremaean Province.

Lectotypification of two Western Australian species of Micromyrtus (Myrtaceae)

RYE, B.L., Nuytsia 15 (1): 153–154 (2002)

Among the species of Micromyrtus Benth. (Myrtaceae) that Bentham (1867) described in "Flora Australiensis", were two Western Australian species that had previously been recognised as distinct taxa by Ferdinand von Mueller. One had been described as Thryptomene elobata (Mueller 1864) and another had been given the manuscript name T. racemosa F. Muell. The purpose of this short communication is to select a lectotype from among the syntypes of each of these species.

Eucalyptus conglobata subsp. perata (Myrtaceae), a new taxon from southern Western Australia and notes on E. series Rufispermae

BROOKER, M.I.H. AND SLEE, A.V., Nuytsia 15 (2): 157–162 (2004)

A new taxon, Eucalyptus conglobata R. Br. ex Maiden subsp. perata Brooker & Slee, from southern Western Australia is described and illustrated. A distribution map for the typical and the new subspecies is provided. A brief discussion of the large E. series Rufispermae to which the species belongs is given.

New species and lectotypifications of some reticulate-nerved Tephrosia (Fabaceae) from north-west Australia and the genus Paratephrosia re-evaluated

COWIE, I.D., Nuytsia 15 (2): 163–186 (2004)

Eight new species of reticulate-nerved Tephrosia are described from the Kimberley Region of Western Australia and the Northern Territory (T. andrewii Cowie, Tephrosia bifacialis Cowie, T. carriemichelliae Cowie, T. ephippioides Cowie, T. gyropoda Cowie, T. humifusa Cowie, T. procera Cowie, and T. valleculata Cowie). Paratephrosia is reduced to synonymy under Tephrosia and a new name, Tephrosia lasiochlaena Cowie, is published for P. lanata. Lectotypifications are provided for six reticulate-nerved and allied taxa of Tephrosia (T. flammea F. Muell. ex Benth., T. nematophylla F. Muell., T. oblonga R. Br. ex Benth., T. polyzyga F. Muell. ex Benth., T. porrecta R. Br. ex Benth. and T. reticulata R. Br. ex Benth.).

Grevillea bipinnatifida subsp. pagna (Proteaceae), a new subspecies from south-west Western Australia

CRANFIELD, R.J., Nuytsia 15 (2): 187–192 (2004)

Restricted to the Swan Coastal Plain of Western Australia, Grevillea bipinnatifida subsp. pagna Cranfield is described. Leaf variation within Grevillea bipinnatifida is also illustrated.

Lichen Census of Western Australia

CRANFIELD, R.J., Nuytsia 15 (2): 193–220 (2004)

A list of known lichen taxa for Western Australia is presented; family, genus, species and infra species are listed with authors and reference to original publication. Biogeographic regions are indicated for each species based on the collections of the Western Australian Herbarium. A table giving statistical data for families and genera is included.

Two new species of Ptilotus (Amaranthaceae) from Western Australia

DAVIS, R.W., Nuytsia 15 (2): 221–226 (2004)

Two new species of Ptilotus (Amaranthaceae) from the central and southern regions of Western Australia are described: Ptilotus chrysocomus R.W. Davis, known from one location near the Carnarvon Range in the vicinity of the abandoned Blue Hill Station, Little Sandy Desert; and Ptilotus halophilus R.W. Davis, known from the margins of Lake Altham and from Lake Hurlstone Reserve in the Lake Grace to Pingrup and Holt Rock to Hyden areas.

Eucalyptus virginea and E. relicta (Myrtaceae), two new rare forest trees from south-western Australia allied to E. lane-poolei, and a new phantom hybrid

HOPPER, STEPHEN D. AND WARDELL-JOHNSON, G., Nuytsia 15 (2): 227–240 (2004)

Eucalyptus virginea and E. relicta are described from Mt Lindesay National Park and from the Whicher Range south-east of Busselton respectively. Both species, together with E. lane-poolei, are in the earliest branching clade of E. ser. Curviptera, which contains many horticulturally attractive large-flowered mallees from semi-arid and desert regions such as E. macrocarpa and E. youngiana. The discovery and description of these two rare relictual forest trees so recently highlights the need for ongoing botanical survey of south-western Australia’s forests. A phantom hybrid, Eucalyptus lane-poolei × relicta, is also described for the first time. It occurs close to populations of E. relicta in the Whicher Range but 150 km south of the nearest recorded E. lane-poolei.

A new species of Bulbine (Asphodelaceae) from Western Australia

KEIGHERY, G.J., Nuytsia 15 (2): 241–244 (2004)

A new species of Bulbine Wolf, B. pendula Keighery, is described and illustrated. The species is confined to cracking clay wetlands in the Pilbara, Carnarvon, Ashburton and Great Sandy Desert Biogeographical Regions of arid Western Australia.

A taxonomic review of the genus Agrostocrinum (Phormiaceae)

KEIGHERY, G.J., Nuytsia 15 (2): 245–252 (2004)

The genus Agrostocrinum F. Muell. is reviewed. Two species are recognised, A. hirsutum (Lindl.) Keighery and A. scabrum (R. Br.) Baillon, both confined to southern Western Australia. A distinctive inbreeding maritime race of the latter, found in scattered populations on granites along the south coast, is described as a new subspecies, A. scabrum subsp. littorale Keighery.

Taxonomic notes on the Angianthus drummondii complex (Asteraceae : Gnaphaliinae)

KEIGHERY, G.J., Nuytsia 15 (2): 253–260 (2004)

Angianthus drummondii is revised and shown to be a species complex of three allopatric species. The complex comprises A. drummondii (Turcz.) Benth., an erect plant with long inflorescence bracts confined to fresh water clay-based wetlands; A. platycephalus Benth., a prostrate plant with short broad inflorescence bracts, growing in fresh to brackish clay soils, is resurrected; and a new species A. halophilus Keighery, an erect plant with short inflorescence bracts, found on gypsum soils, is described.

Taxonomy of the Calytrix ecalycata complex (Myrtaceae)

KEIGHERY, G.J., Nuytsia 15 (2): 261–268 (2004)

The part of the genus Calytrix Labill. that was formerly regarded as the monotypic genus Calythropsis C.A. Gardner is shown to be composed of two allopatric species, Calytrix ecalycata Craven and C. pimeleoides C.A. Gardner ex Keighery. The former species is comprised of three geographically separated subspecies, C. ecalycata subsp. ecalycata, C. ecalycata subsp. brevis Keighery and C. ecalycata subsp. pubescens Keighery.

Two new species of Dampiera (Goodeniaceae) from the Pilbara region, Western Australia

LEPSCHI, B.J., TRUDGEN, MALCOLM E. AND VAN LEEUWEN, S., Nuytsia 15 (2): 269–276 (2004)

Dampiera anonyma and Dampiera metallorum Lepschi & Trudgen, two geographically restricted taxa from the Pilbara region of Western Australia, are described, illustrated and their distributions mapped.

A review of Hibbertia hemignosta and its allies (Dilleniaceae) from Western Australia

WHEELER, J.R., Nuytsia 15 (2): 277–298 (2004)

The group of species that includes Hibbertia hemignosta (Steud.) J.R. Wheeler is reviewed and a key provided. The new species Hibbertia acrotrichion J.R. Wheeler and Hibbertia chartacea are described. Two new varieties of Hibbertia hibbertioides (Steud.) J.R. Wheeler, var. meridionalis and var. pedunculata, and two new varieties of Hibbertia pulchra Ostenf., var. acutibractea and var. crassinervia, are also described. All taxa are mapped and illustrated. Both new species have conservation priority, but the four new varieties are not considered endangered.

Miscellaneous new Hibbertia species (Dilleniaceae) from the south coast and adjacent interior of Western Australia

WHEELER, J.R., Nuytsia 15 (2): 299–310 (2004)

Five new species of Hibbertia Andr. are formally described. Hibbertia oligantha J.R. Wheeler, Hibbertia pachyphylla J.R. Wheeler, Hibbertia papillata J.R. Wheeler, Hibbertia psilocarpa J.R. Wheeler and Hibbertia turleyana J.R. Wheeler all from the south coast area of the South-West Botanical Province (Esperance Plains region and Mallee region) with two species extending to the southern Eremaean Province (Coolgardie region). All species are mapped and the three having conservation priority are illustrated.

An interim key to the Western Australian species of Hibbertia (Dilleniaceae)

WHEELER, J.R., Nuytsia 15 (2): 311–320 (2004)

A key is presented to the Western Australian species of Hibbertia following the formal recognition of numerous new species in recent years.

Recognition of Eucalyptus quaerenda at specific rank

BYRNE, M., Nuytsia 15 (2): 321–323 (2004)

Phylogenetic analysis of the relationships between the taxa in the Eucalyptus angustissima complex have been investigated using RFLP analysis of nuclear genome (Elliott and Byrne 2003) (Voucher specimens at PERTH: W. O’Sullivan 127, 357, 812, 968, 966, 1183, 1190, 1193, 1194). This study assessed populations of E. angustissima subsp. angustissima, E. angustissima subsp. quaerenda, E. foliosa and E. misella and found all four taxa to be genetically distinct. Eucalyptus angustissima was not monophyletic as the two subspecies did not cluster together. Eucalyptus misella was sister species to E. angustissima subsp. angustissima, and E. foliosa was sister species to both these taxa. Eucalyptus angustissima subsp. quaerenda was the most distinct of the four taxa.

Correction of type locality and flowering time for Melaleuca apostiba (Myrtaceae)

LEWINGTON, M.A., Nuytsia 15 (2): 324–324 (2004)

Melaleuca apostiba K.J. Cowley was described from a specimen collected by R. Morland during a Lands & Surveys Department expedition to the Officer Basin in 1963. The holotype of M. apostiba has the collecting number M10, with the label recording the locality as ‘between Carnegie and Giles’ and the collection date as June 1963.

A first report of Leptochloa panicea subsp. brachiata (Poaceae) from Western Australia

SNOW, NEIL, Nuytsia 15 (2): 325–326 (2004)

The genus Leptochloa P. Beauv. (Poaceae) is represented by nine species in Australia (Sharp and Simon 2002). Three of these include subspecies, bringing the total number of taxa in Australia to fourteen. I report here the first confirmed occurrence in Western Australia of Leptochloa panicea (Retz.) Ohwi subsp. brachiata (Steud.) N. Snow, a non-native taxon of neotropical origin.

Nomenclatural notes on two species of Hibbertia (Dilleniaceae)

WHEELER, J.R., Nuytsia 15 (2): 327–328 (2004)

Examination of type material of Hibbertia rhadinopoda and closely related taxa has revealed that there is an earlier name for H. rhadinopoda in the genus Pleurandra Labill. The necessary new combination is made here. A lectotype is also chosen for Hibbertia inclusa.

Two new species of Brachyloma (Epacridaceae) from the South West Botanical Province of Western Australia

CRANFIELD, R.J., Nuytsia 15 (3): 331–336 (2005)

Endemic to the South West Botanical Province of Western Australia, Brachyloma delbi Cranfield and Brachyloma mogin Cranfield are described, illustrated and mapped.

Further new taxa in Dryandra R.Br. (Proteaceae : Grevilleoideae)

GEORGE, A.S., Nuytsia 15 (3): 337–346 (2005)

The following new taxa in Dryandra are described: D. prionotes A.S. George, D. ferruginea subsp. magna A.S. George, D. fraseri var. crebra A.S. George, D. fraseri var. effusa A.S. George, D. ionthocarpa subsp. chrysophoenix A.S. George and D. pteridifolia subsp. inretita A.S. George.

Reinstatement of Burchardia congesta (Colchicaceae)

KEIGHERY, G.J. AND MUIR, W., Nuytsia 15 (3): 347–354 (2005)

Burchardia R. Br. is an endemic Australian genus of five species in the Colchicaceae. Burchardia umbellata R. Br. is a widespread taxon occurring disjunctly in temperate Eastern Australia and in south-west Western Australia. Studies on the morphology, ecology and anatomy of the Western and Eastern populations have shown marked discontinuities in character sets, leading to the recognition that each area is a seperate taxon. The name B. congesta Lindl. is re-instated for the Western populations and the name B. umbellata restricted to the Eastern populations.

A taxonomic revision of Drosera section Stolonifera (Droseraceae) from south-west Western Australia

LOWRIE, A., Nuytsia 15 (3): 355–394 (2005)

Ten species are recognized in the Drosera section Stolonifera: D. fimbriata DeBuhr, D. humilis Planchon, D. platypoda Turcz., D. porrecta Lehm., D. purpurascens Schlott., D. ramellosa Lehm., and D. stolonifera Endl., including three new combinations: D. monticola (Lowrie & Marchant) Lowrie, D. prostrata (Marchant & Lowrie) Lowrie and D. rupicola (Marchant) Lowrie. All are endemic to the south-west of Western Australia and belong in Drosera L. subgen. Ergaleium DC. sect. Stolonifera DeBuhr. Each taxon is described in detail as well as illustrated. A key is provided to all taxa in sect. Stolonifera. A schematic illustration of all ten species life-form is provided for comparison and cross referencing with each other. SEM micrographs of the seeds of each species, field and cultivation studies and historical investigations are also presented.

A rare and endangered new subspecies of Eucalyptus sargentii (Myrtaceae) with high potential for revegetation of saline sites from south-western Australia and notes on E. diminuta and E. sargentii subsp. fallens

NICOLLE, D., Nuytsia 15 (3): 395–402 (2005)

Eucalyptus sargentii subsp. onesia Nicolle subsp. nov. is described. It is known from less than ten small populations in the central wheatbelt of southwest Western Australia, with all populations occurring on highly saline sites and possibly endangered by increasing salinisation. E. sargentii subsp. onesia differs from subsp. sargentii primarily in the lignotuberous, mallee habit, making it preferable to subsp. sargentii for salt reclamation projects, due to its propensity to regenerate following fire or cutting. Extensive field examination, seedling trials and observations of cultivated material have indicated that E. sargentii subsp. fallens is not specifically distinct from E. diminuta and I regard the former as synonymous with the latter. E. diminuta is recircumscribed to include populations from Eurardy Station in the north to near Watheroo in the south.

Re-assessment of the saline-dwelling Eucalyptus spathulata complex (Myrtaceae) from southern Western Australia

NICOLLE, D. AND BROOKER, M.I.H., Nuytsia 15 (3): 403–430 (2005)

The circumscription of taxa within the Eucalyptus spathulata complex is revised. The status of E. vegrandis is resolved and the new species E. orthostemon Nicolle & Brooker is described to accommodate populations of mallees previously and erroneously referred to E. vegrandis. The new taxon E. vegrandis subsp. recondita Nicolle & Brooker is described to accommodate coarse, relatively broad-leaved mallees in the Stirling Range area, previously referred under the manuscript name E. ‘recondita’. E. suggrandis is recircumscribed, with subsp. alipes (as to the type – the mallet/obligate seeder variant) raised to specific status as E. alipes (L. Johnson & K. Hill) Nicolle & Brooker. E. mimica is recircumscribed as an obligate seeder (mallet) species and E. suggrandis subsp. promiscua Nicolle and Brooker described from the Lake Grace – Pingrup area, previously being confused with both E. mimica and E. suggrandis. The new subspecies, E. spathulata subsp. salina Nicolle & Brooker, is described from mallets restricted to the Salt River drainage system of the northern part of the southern wheatbelt. E. cernua is recircumscribed as an obligate seeder (mallet) species with resprouter (mallee) populations, previously included under E. cernua, published as E. proxima Nicolle & Brooker. The manuscript name E. ‘verruculosa’ is considered to be E. suggrandis subsp. suggrandis, as to the proposed type. A key and distribution maps for the E. spathulata complex are provided.

New taxa, a new record and a rediscovery in Western Australian Haloragis (Haloragaceae)

ORCHARD, A.E., LEPSCHI, B.J. AND HISLOP, M., Nuytsia 15 (3): 431–444 (2005)

Variation and distribution in taxa of the Haloragis gosseiH. trigonocarpa group is discussed and two new taxa, H. maierae Orchard and H. gossei var. inflata Orchard are described. Rediscovery of H. platycarpa is noted, and an amended description of this rare species is provided. Variation in the H. aculeataH. scoparia group is discussed, and a widely disjunct new record, H. glauca forma glauca, is noted for the State.

A taxonomic review of Dicrastylis sect. Corymbosae (Lamiaceae: Chloantheae), incorporating Mallophora as a new synonym

RYE, B.L., Nuytsia 15 (3): 445–456 (2005)

Following a recent recommendation to conserve the name Dicrastylis Drumm. ex Harv. over Mallophora Endl., the two species previously included in the latter genus are here transferred to Dicrastylis sect. Corymbosae Munir. To achieve this, two new combinations, Dicrastylis globiflora (Endl.) Rye and D. rugosifolia (Munir) Rye, are made. The circumscription of Dicrastylis sect.Corymbosae is further altered by the removal of D. nicholasii F. Muell., and D. glauca is reduced to a synonym of D. corymbosa. A review of the section is presented. It is now comprised of five white-flowered species with cymes condensed into corymbosely arranged clusters, and is restricted to the south-west of Western Australia.

A taxonomic update of Petrophile sect. Arthrostigma (Proteaceae)

RYE, B.L. AND HISLOP, M., Nuytsia 15 (3): 457–484 (2005)

Petrophile sect. Arthrostigma (Endl.) Benth. is described, P. teretifolia R. Br. is chosen as its lectotype, and a key is given for all members of the section. Petrophile filifolia R. Br. and P. juncifolia Lindl. are reinstated. Descriptions are also given for five new taxa, Petrophile filifolia subsp. laxa Rye & Hislop, P. pilostyla, P. pilostyla subsp. austrina, P. pilostyla subsp. syntoma and P. prostrata, and for two species that are now more narrowly circumscribed, P. brevifolia R. Br. and P. longifolia R. Br. A lectotype is also selected for P. longifolia. Pollen presenter characters, which are of particular importance in this group, are compared in a table and are illustrated for all of the named taxa.

A new heterocarpidic fruit type for the Myrtaceae, with dehiscent and indehiscent loculi, in two genera from Western Australia

RYE, B.L. AND TRUDGEN, MALCOLM E., Nuytsia 15 (3): 485–494 (2005)

A form of heterocarpidy with dehiscent and indehiscent loculi present in the same fruit is described from species in two genera of Myrtaceae occurring in the south-west of Western Australia. This extreme development of heterocarpidy results in a very unusual fruit type, one that has not previously been described for the Myrtaceae. It is also apparently rare in the angiosperms, although smaller differences between carpels (mostly in their size and the number of seeds) are not uncommon. In Astus Trudgen & Rye and the Baeckea robusta F. Muell. complex, two types of ovary loculi develop, with one type dehiscent by a suture on the floral disc and the other type lower in the ovary and indehiscent. The occurrence of this heterocarpidic fruit type in two fairly different genera is considered to be a convergent development rather than indicating a particularly close relationship. The heterocarpidic fruit in these groups is described and illustrated, and its adaptive and taxonomic significance discussed.

Astus, a new Western Australian genus of Myrtaceae with heterocarpidic fruits

TRUDGEN, MALCOLM E. AND RYE, B.L., Nuytsia 15 (3): 495–512 (2005)

Astus Trudgen & Rye, a new genus of Myrtaceae with four species endemic to the south-west of Western Australia, is described and its affinities discussed. While the genus is circumscribed on the basis of a range of characteristics that separate it from related genera, all its members are unusual in having a fruit with both dehiscent and indehiscent loculi. In addition, two of the species have a marked heteromorphy in their calyx lobes. Astus is one of a number of genera belonging to the tribe Chamelaucieae that have reniform seeds. While the floral morphology of the new genus does not readily indicate which of the other reniform-seeded genera it is closest to, DNA evidence indicates affinities to the Eastern Australian genus Triplarina Raf. The type species, Astus tetragonus (F. Muell. ex Benth.) Trudgen & Rye, is based on Baeckea tetragona F. Muell. ex Benth. Astus duomilius Trudgen & Rye, A. subroseus Trudgen & Rye and A. wittweri Trudgen & Rye are new species. A key to the species, descriptions, illustrations and distribution maps are provided.

Goodenia pedicellata (Goodeniaceae), a new species from the Pilbara bioregion of Western Australia

SAGE, L.W. AND DIXON, K.W., Nuytsia 15 (3): 513–516 (2005)

A new species of conservation priority, Goodenia pedicellata, is described, illustrated, mapped and compared with G. cusackiana (F. Muell.) Carolin. Surveys are needed to accurately determine the rarity of the species in the wild, currently known only from the holotype location.

Rulingia borealis, a new combination based on R. malvifolia var. borealis (Malvaceae s.l. or Sterculiaceae)

WILKINS, C.F., Nuytsia 15 (3): 517–522 (2005)

Rulingia malvifolia Steetz is an illegitimate name since Steetz, in describing R. malvifolia cited Commersonia cygnorum Steud. in synonomy. Pritzel (1901) then described Rulingia malvifolia var. borealis E.Pritz. This variety is here raised to species rank to become R. borealis (E.Pritz.) C.F.Wilkins.

New and noteworthy plant species recognised as naturalised in Western Australia

KEIGHERY, G.J., Nuytsia 15 (3): 523–528 (2005)

The format of this paper follows that of Heenan et al. (2002) for New Zealand and Hosking et al. (2003) for New South Wales. Species are grouped under Monocotyledons or Dicotyledons, then listed aphabetically by family and scientific name, common name (when available), the location of a taxon description, natural region where the weed has been recorded following the Interim Biogeographic Regionalisation for Australia (Thackway & Cresswell 1995), habitats, first records and area of origin.